Publications and communications

You can find here my publications, my PhD thesis and some communications. If you are interested in one of my paper, please send me an e-mail at

Publications in indexed journals

.Dehon, M., Perrard, A., Engel, M.S., Nel, A. & Michez, D. 2017. Antiquity of cleptoparasitism among bees revealed by morphometric and phylogenetic analysis of a Paleocene fossil nomadine (Hymenoptera: Apidae). Systematic Entomology. Sous Presse.

.Perrard, A., Grimaldi, D. & Carpenter, J.M. 2017. Early lineages of Vespidae (Hymenoptera) in Cretaceous amber. Systematic Entomology 42: 379–386. (PDF)

.Perrard, A., Lopez-Osorio, F. & Carpenter, J.M. 2016. Phylogeny, landmark analysis and the use of wing venation to study the evolution of social wasps (Hymenoptera: Vespidae: Vespinae). Cladistics In Press. DOI:10.1111/cla.12138. (PDF)

. Carpenter, J.M., Kojima, J.-I., Dvorak, L. & Perrard, A. 2015. Taxonomic notes on Vespinae (Hymenoptera: Vespidae). Entomologica Americana, 121: 35–37. (PDF)

.Lopez-Osorio, F., Perrard, A., Pickett, K.M., Carpenter, J.M. & Agnarsson, I. 2015. Phylogenetic tests reject Emery’s rule in the evolution of social parasitism in yellowjackets and hornets (Hymenoptera: Vespidae, Vespinae). Royal Society Open Science 2:150159. DOI:10.1098/rsos.150159 (PDF)

.Perrard, A. & Loope, K.J. 2015. Patriline differences reveal genetic influence on forewing size and shape in a Yellowjacket wasp (Hymenoptera: Vespidae: Vespula flavopilosa Jacobson, 1978) PLoS ONE 10(7): e0130064. (PDF)

.Arca, M., Mougel, F., Guillemaud, T., Dupas, S., Rome, Q., Perrard, A., Muller, F., Fossoud, A., Capdevielle-Dulac, C., Torres-Leguizamon, M., Chen, X.X., Tan, J.L., Jung, C., Villemant, C., Arnold & G., Silvain, J.-F. 2015. Reconstructing the invasion and the demographic history of the Yellow-legged hornet, Vespa velutina, in Europe. Biological Invasions 17(8): 2357–2371. DOI : 10.1007/s10530-015-0880-9. (PDF)

.Rome, Q., Muller, F. J., Touret-Alby, A., Darrouzet, E., Perrard, A. and Villemant, C. 2015. Caste differentiation and seasonal changes in Vespa velutina (Hym.: Vespidae) colonies in its introduced range. Journal of Applied Entomology DOI:10.1111/jen.12210. (PDF)

.Perrard, A., Baylac, M., Carpenter, J.M., & Villemant, C. 2014. Evolution of wing shape in hornets: why is the wing venation efficient for species identification? Journal of Evolutionary Biology 27: 2665–2675. (PDF)

. Perrard, A., Arca, M., Rome, Q., Muller, F., Tan, J., Bista, S., Nugroho, H., Baudoin, R., Baylac, M., Silvain, J.-F., Carpenter, J.M., & Villemant, C. 2014 Geographic variation of melanisation patterns in a hornet species: genetic differences, climatic pressures or aposematic constraints? Plos ONE 9: e94162 (PDF)

. Perrard, A., Nel, A., & Carpenter, J.M. 2014 A new paper wasp from Late Eocene of France (Hymenoptera: Vespidae: Polistinae). Zootaxa 3753: 542-548. (PDF)

. Fabre, A.C., Cornette, R., Perrard, A., Boyer, D., Prasad, G.V.R., Hooker, J.J., & Goswami, A. 2014 A three-dimensional morphometric analysis of the locomotory ecology of Deccanolestes, a eutherian mammal from the late Cretaceous of India. Journal of Vertebrate Paleontology 34: 146-156. (PDF)

. Perrard, A., Pickett, K.M., Villemant, C., Kojima, J.-I., & Carpenter, J.M. 2013. Phylogeny of the genus Vespa: a total evidence approach (Hymenoptera: Vespidae; Vespinae, Vespa). Journal of Hymenoptera Research 32: 1-15 (PDF). 

. Barbet-Massin, M., Rome, Q., Muller, F., Perrard, A., Villemant, C. & Jiguet, F. 2013. Climate change increases the risk of invasion by the Yellow-legged hornet. Biological Conservation 157: 4 – 10. (PDF)

. Fontaine, B., Perrard, A., & Bouchet, P. 2012. 21 Years between Discovery and Description: the Shelf Life of New Species Reveals Taxon, Social and Geopolitical Biases. Current Biology 22: 943 – 944. (PDF)

. Perrard, A., Villemant, C., Carpenter, J.M., & Baylac, M. 2012. Differences in caste dimorphism among three hornet species (Hymenoptera: Vespidae): forewing size, shape and allometry. Journal of Evolutionary Biology 25: 1389-1398. (PDF)

. Carpenter, J. M., Dvorák, L., Kojima, J. I., Nguyen, L. T. P., Perrard, A., & Pickett, K. M. 2011. Taxonomic Notes on the Vespinae of Yunnan (Hymenoptera: Vespidae). American Museum Novitates 3709: 1–10. (PDF)

. Villemant, C., Barbet-Massin, M., Perrard, A., Muller, F., Gargominy, O., Jiguet, F., & Rome, Q. 2011. Predicting the invasion risk by the alien bee-hawking Yellow-legged hornet Vespa velutina nigrithorax across Europe and other continents with niche models. Biological Conservation 144 (9): 2142-2150. (PDF)

. Perrard, A., Haxaire, J., Rortais, A., & Villemant, C. 2009. Observations on the colony activity of the Asian hornet Vespa velutina Lepeletier 1836 (Hymenoptera: Vespidae: Vespinae) in France. Annales de la Société Entomologique de France 45 (1): 119-127. (PDF)

Book chapters

. Villemant, C., Muller, F., Rome, Q., Perrard, A., Barbet-Massin, M., & Jiguet, F. 2014. Estimating potential range expansion and environmental impact of the invasive bee hawking hornet, Vespa velutina nigrithorax. In: Devillers J. (Ed.), In Silico Bees. CRC Press, Boca Raton, Florida.

Publication in other peer-reviewed journals

. Perrard, A., Muller, F., Rome, Q., & Villemant, C. 2011. Observations on the yellow-legged hornet Vespa velutina Lepeletier, 1836 (Hymenoptera, Vespidae). Bulletin de la Société Entomologique de France 116 (2): 159-164. (PDF)

. Rome, Q., Perrard, A., Muller, F., & Villemant, C. 2011. Monitoring and control modalities of a honeybee predator, the yellow-legged hornet Vespa velutina nigrithorax (Hymenoptera: Vespidae). Aliens: The Invasive Species Bulletin 31: 7-15. (PDF)

. Villemant, C., Muller, F., Haubois, S., Perrard, A., Darrouzet, E., & Rome, Q. 2011. Bilan des travaux (MNHN et IRBI) sur l’invasion en France de Vespa velutina, le frelon asiatique prédateur d’abeilles. In Proceedings of Journée Scientifique Apicole, Arles 11: 3–12. (PDF)

PhD Thesis

Perrard, A. 2012. Systematique et Morphometrie géométrique : l’évolution de la nervation alaire au sein du genre Vespa (Hyménoptères : Vespidés) (PDF with english summary for each chapter)

[Systematics & geometric morphometrics: evolution of the wing venation within the genus Vespa (Hymenoptera: Vespidae)]


. Perrard, A. Communication: Phylomorphospace: Combining morphospace and phylogeny. Invited lecturer at the morphometrics workshop of Botany 2015, Edmonton, Canada. July, 2015.

. Perrard, A. Lopez-Osorio, F., Carpenter J.M. Communication: Combining landmark data and phylogenetic analyses: Can we tell the story of wasps from the lines of their wings? RGGS comparative biology seminar, New York, USA. March, 2015.

. Perrard, A. Lopez-Osorio, F., Carpenter J.M. Communication: Evolution of Yellow-jackets and Hornets (Vespidae: Vespinae) and the use of wing morphometrics in phylogenetics. International Society of Hymenopterists meeting, Cusco, Peru. August, 2014.

. Perrard, A. Communication: Analysis of the genetic diversity and colour variation of the Asian populations of the hornet Vespa velutina. Systematics and evolution seminar, Paris, France. April 2014.

. Perrard, A. Communication:  Evolution of the wing shape in social wasps: morphometrics and phylogenies. Invited lecturer at the Morphometrics workshop in Naturalis, Leiden, The Netherlands. September 2013.

. Perrard, A. Communication: Mélanges gaussiens et signal phylogénétique: variation de l’aile chez les Vespidés. Seminaire Morpho, Paris, France. July 2013.

. Perrard, A., Baylac, M., Carpenter, J.M., Villemant, C. Communication: Wing shape evolution in social wasps and phylogenetic signal. MEETING EVOLUTION 2013, Snowbird, Utah, USA. June 2013.

. Perrard, A., Debat V., Muller F., Rome Q., Carpenter J.M., Baylac M., Villemant, C. Lightning talk: Evolution of the wing shape in insects: drift and selection in the Yellow-Legged Hornet Vespa velutina (Hymenoptera: Vespidae). MEETING EVOLUTION 2013, Snowbird, Utah, USA. June 2013.

. Perrard, A. Communication: Mclust: Déceler des groupes dans un jeu de données grâce aux mélanges gaussiens. SeminR, Paris, France. July 2013.

. Perrard, A., Villemant, C., Carpenter, J.M., Baylac, M. Communication: Caste dimorphism in hornets: size, shape and allometry of the wing. VIIème Seminaire de Morphométrie et Evolution des Formes, Lyon, France. May 2012.

. Perrard, A. Poster: Sociality in Hornets: Are queens big workers? Paris Interdisciplinary PhD Symposium, Paris, France. June 2011.

. Perrard, A., Rome, Q., Muller, F., Villemant, C. Communication: The Yellow-Legged Hornet Vespa velutina (Hym.: Vespidae): a new invader in France. IXth European Congress of Entomology, Budapest, Hungary. August 2010.

. Perrard, A., Carpenter, J.M., Pickett, K.M., Villemant, C. Poster: Phylogénie du genre Vespa (Hyménoptère : Vespidae) sur la base de données morphologiques et moléculaires. Journées de la Société Française de Systématique, Anger, France. October 2009.


One response to “Publications and communications

  1. rosa

    muchisimas gracias.
    la website esta muy bien documentada y es de gran ayuda
    un saludo

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